Dr. Holmes completed his undergraduate degree at Harvard College in 1959, his medical degree at Cornell University Medical College in 1963 and received his Ph.D. in Microbiology from the University of Hawaii in 1967. Dr. Holmes has been a member of the University of Washington faculty since 1969, holding educational leadership positions in areas such as medicine, microbiology, epidemiology and global health. He is the principal investigator for the International Training & Education Center on Health (I-TECH), a collaborative program between the University of Washington and the University of California, one of the largest HIV/AIDS training programs in the world. Dr. Holmes also directs the University of Washington/Fred Hutchinson Cancer Research Center’s Center for AIDS Research. He has participated in research on STDs for over 40 years in Africa, Latin America, Southeast Asia, and the Western Pacific. He has authored over 550 peer-reviewed publications and edited 30 books, monographs, and journal supplements.